* Corresponding authors

^a Single-Cell Center, CAS Key Laboratory of Biofuels and Shandong Key Laboratory of Energy Genetics, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, China
E-mail: xujian@qibebt.ac.cn, mabo@qibebt.ac.cn

^b College of Chemical Science and Engineering, Qingdao University, Qingdao, China

^c Department of Engineering Science, University of Oxford, Parks Road, Oxford OX1 3PJ, UK

^d Public Laboratory and CAS Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, China
E-mail: yangml@qibebt.ac.cn