* Corresponding authors

^a Department of Bioengineering, University of Pennsylvania, Philadelphia, PA 19104, USA
E-mail: mhchoi1@seas.upenn.edu, atimperm@seas.upenn.edu

^b Department of Biochemistry and Biophysics, University of Pennsylvania, Philadelphia, PA, USA

^c Department of Mechanical Science and Engineering, University of Illinois at Urbana-Champaign, Urbana, IL 61801, USA

^d Department of Physics, Utah State University, Logan, UT 84322, USA