* Corresponding authors

^a Integrative Science Center of Germplasm Creation in Western China (CHONGQING) Science City and Southwest University, College of Agronomy and Biotechnology, Southwest University, Chongqing, P.R. China
E-mail: shahidbiochem@wmc.edu.pk, swu0120@swu.edu.cn, qmc2020@email.swu.edu.cn

^b Engineering Research Center of South Upland Agriculture, Ministry of Education, Chongqing, P.R. China
E-mail: drlukun@swu.edu.cn

^c Department of Botany, University of Science and Technology Bannu, KPK, Pakistan
E-mail: sirabkhan246@gmail.com

^d Institute of Tropical Bioscience and Biotechnology, Chinese Academy of Tropical Agricultural Sciences, Haikou-571101, China
E-mail: talhajaved54321@gmail.com

^e Department of Agronomy, Abdul Wali Khan University Mardan, Khyber Pakhtunkhwa 23200, Pakistan
E-mail: shahfahad@awkum.edu.pk

^f Department of Natural Sciences, Lebanese American University, Byblos, Lebanon